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Fridley, J.D., Bellingham, P., Closset-Kopp, D., Daehler, C., Dechoum, M., Martin, P., Murphy, H., Rojas-Sandoval, J. & Tng, D. 2023. A general hypothesis of forest invasions by woody plants based on whole-plant carbon economics. Journal of Ecology 111: 4-22. [online] [PDF]

Fridley, J.D. and Palmquist, K.A. 2023. Species richness in herbaceous vegetation. In: Encyclopedia of Biodiversity, 3rd Edition. Elsevier.

Lynn, J.S., Fridley, J.D. & Vandvik, V. 2023. More than what they eat: uncoupled biophysical constraints underlie geographic patterns of herbivory. Ecography 2023: e06114. [online] [PDF]


Fridley, J.D., Bauerle, T., Craddock, A., Ebert, A., Frank, D., Heberling, J.M., Hinman, E., Jo, I., Martinez, K., Smith, M., Woolhiser, L. & Yin, J. 2022. Fast but steady: an integrated leaf-stem-root trait syndrome for woody forest invaders. Ecology Letters 25: 900-912. [online]

Beslity, J., Shaw, S.B., Drake, J.E., Fridley, J.D., Stella, J.C., Stark, J. & Singh, K. 2022. A low cost, low power sap flux device for distributed and intensive monitoring of tree transpiration. HardwareX: e00351. [online]

Stark, J.M. and Fridley, J.D. 2022. Microclimate-based species distribution models in complex forested terrain indicate widespread cryptic refugia under climate change. Global Ecology and Biogeography 31: 562-575. [online] [highlighted in The Hellbender Press, April 2023]

Lamit, L., Giovati, A., Jo, I., Frank, D.A. and Fridley, JD. 2022. Woody invaders are more highly colonized by arbuscular mycorrhizal fungi than congeneric native species in a common garden. American Journal of Botany 109: 655-663. [PDF]


In memoriam: Phil Grime, 1935-2021 [Trends in Ecology & Evolution] [PDF]

Pierce, S. and Fridley, J.D. 2021. John Philip Grime. Biographical Memoirs of Fellows of the Royal Society 71: 251-270. [online]

Fridley, J.D., Jo, I., Hulme, P.E. & Duncan, R.P. 2021. A habitat-based assessment of the role of competition in plant invasions. Journal of Ecology, in press. [online]

Griffin-Nolan, R.J., Mohanbabu, N., Araldi-Brondolo, S., Ebert, A.R., LeVonne, J., Lumbsden-Pinto, J.I., Roden, H., Stark, J.R., Tourville, J., Becklin, K.M., Drake, J.E., Frank, D.A., Lamit, L. J. and Fridley, J.D. 2021. Friend or foe? The role of biotic agents in drought-induced plant mortality. Plant Ecology 222: 537-548. [online]

Brandt, A.J., Bellingham, P.J., Duncan, R.P., Etherington, T.R., Fridley, J.D., Howell, C.J., Hulme, P.E., Jo, I., McGlone, M.S., Richardson, S.J., Sullivan, J.J., Williams, P.A. & Peltzer, D.A. 2021. Naturalised plants transform the composition and function of the New Zealand flora. Biological Invasions 23: 351-366. [online] [PDF]


Hull, V., Frank, D.A. and Fridley, J.D. 2020. Woody invaders do not alter rhizosphere microbial activity in a temperate deciduous forest. Biological Invasions 22: 2599-2608. [online] [PDF]

Hinman, E.D. and Fridley, J.D. 2020. Impacts of experimental defoliation on native and invasive saplings: are native species more resilient to canopy disturbance? Tree Physiology 40: 969-979. [online] [PDF]

Jo, I., Fridley, J.D. and Frank, D.A. 2020. Rapid leaf litter decomposition of deciduous understory shrubs and lianas mediated by mesofauna. Plant Ecology 221: 63-68. [PDF]


Kong, D. and Fridley, J.D. 2019. Does plant biomass partitioning reflect energetic investments in carbon and nutrient foraging? Functional Ecology 33: 1627-1637. [PDF]

Martinez, K.A., Fridley, J.D., Oguchi, R., Aiba, M. and Hikosaka, K. 2019. Functional shifts in leaves of woody invaders of deciduous forests between their home and away ranges. Tree Physiology 39: 1551-1560. [PDF]

Heberling, J.M., McDonough MacKenzie, C., Fridley, J.D., Kalisz, S., and Primack, R. 2019. Phenological mismatch with trees reduces wildflower carbon budgets. Ecology Letters 222: 616-623. [PDF]

Hinman, E.D., Fridley, J.D., and Parry, D. 2019. Plant defense against generalist herbivores in the forest understory: a phylogenetic comparison of native and invasive species. Biological Invasions 21: 1269-1281. [PDF]

Heberling, J.M., Cassidy, S.T., Fridley, J.D., and Kalisz, S. 2019. Carbon gain phenologies of spring-flowering perennials in a deciduous forest indicate a novel niche for a widespread invader. New Phytologist 221: 778-788. [PDF]

Lynn, J.S. and Fridley, J.D. 2019. Geographic patterns of plant-herbivore interactions are driven by soil fertility. Journal of Plant Ecology 12: 653-661. [PDF]


Fridley, J.D. and Wright, J.P. 2018. Temperature accelerates the rate fields become forests. Proceedings of the National Academy of Sciences USA 115: 4702-4706. [online] [PDF]

Hinman, E.D. and Fridley, J.D. 2018. To spend or to save? Assessing energetic growth-storage tradeoffs in native and invasive woody plants. Oecologia 188: 659-669. [PDF]

Martinez, K.A. and Fridley, J.D. 2018. Acclimation of leaf traits in seasonal light environments: are non-native species more plastic? Journal of Ecology 106: 2019-2030. [PDF]

Frank, D.A., Wallen, R., Hamilton, B., White, P.J., and Fridley, J.D. 2018. Manipulating the system: how large herbivores control bottom-up regulation of grasslands. Journal of Ecology 106: 434-443. [PDF]

Basto, S., Thompson, K., Grime, J.P., Fridley, J.D., Calhim, S., Askew, A.P. and Rees, M. 2018. Severe effects of long-term drought on calcareous grassland seed banks. npj Climate and Atmospheric Science, 1(1): 1. [PDF]


Fridley, J.D. 2017. Plant energetics and the synthesis of population and ecosystem ecology. Journal of Ecology 105: 95-110. [online] [PDF]

Jo, I., Fridley, J.D. and Frank, D.A. 2017. Invasive plants accelerate nitrogen cycling: evidence from experimental woody monocultures. Journal of Ecology 105: 1105-1110. [online] [PDF]

Sayer, E.J., Oliver, A.E., Fridley, J.D., Askew, A.P., Mills, R.T.E. and Grime, J.P. 2017. Links between soil microbial communities and plant traits in a species-rich grassland under long-term climate change. Ecology & Evolution 7: 855-862. [online] [PDF]

Zohner, C., Benito, B., Fridley, J.D., Svenning, J.-C. and Renner, S. 2017. Spring predictability explains different leaf-out strategies in the woody floras of North America, Europe, and East Asia. Ecology Letters 20: 452-460. [PDF]

Heberling, J.M., Jo, I., Kozhevnikov, A., Lee, H. and Fridley, J.D. 2017. Biotic interchange in the Anthropocene: strong asymmetry in East Asian and Eastern North American plant invasions. Global Ecology and Biogeography 26: 447-458. [PDF]


Fridley, J.D., Lynn, J.S., Grime, J.P. and Askew, A.P. 2016. Longer growing seasons shift grassland vegetation towards more-productive species. Nature Climate Change 6: 865-868. [online] [PDF]

Jo, I., Fridley, J.D. and Frank, D.A. 2016. More of the same? In situ leaf and root decomposition rates do not vary between 80 native and non-native deciduous forest species. New Phytologist 209: 115-122. [PDF] [Commentary article]

Heberling, J.M., Kichey, T., Decocq, G., and Fridley, J.D. 2016. Plant functional shifts in the invaded range: a test with reciprocal forest invaders of Europe and North America. Functional Ecology 30: 875-884. [PDF]

Heberling, J.M. and Fridley, J.D. 2016. Invaders do not require high resource levels to maintain physiological advantages in a temperate deciduous forest. Ecology 97: 874-884. [PDF] 

Negoita, L., Fridley, J.D., Lomolino, M., Mittelhauser, G., Craine, J., and Weiher, E. 2016. Isolation-driven functional assembly of plant communities on islands. Ecography 39: 1066-1077. [online]


Fridley, J.D. and Craddock, A. 2015. Contrasting growth phenology of native and invasive forest shrubs mediated by genome size. New Phytologist 207: 659-668. [PDF] [Research Highlight in Nature PlantsRecommended by Faculty of 1000

Fraser, L.H., et al. 2015. Worldwide evidence of a unimodal relationship between productivity and plant species richness. Science 349: 302-305. [online]

Lesser, M.R. and Fridley, J.D. 2015. Global change at the landscape level: relating regional and landscape-scale drivers of historical climate trends in the Southern Appalachians. International Journal of Climatology 36: 1197-1209. [PDF]

Ravenscroft, C.H., Whitlock, R., and Fridley, J.D. 2015. Rapid genetic divergence in response to 15 years of simulated climate change. Global Change Biology 21: 4165-4176. [PDF]

Frank, D.A., Pontes, A., Maine, E., and Fridley, J.D. 2015. Fine-scale belowground species associations in temperate grassland. Molecular Ecology 24: 3206-3216. [PDF]

Yin, J., Fridley, J.D., Smith, M., and Bauerle, T. 2015. Xylem vessel traits predict the leaf phenology of native and non-native understory species of temperate deciduous forests. Functional Ecology 30: 206-214. [PDF]

Jo, I., Fridley, J.D. and Frank, D.A. 2015. Linking above- and belowground resource use strategies for native and invasive species of temperate deciduous forests. Biological Invasions 17: 1545-1554. [PDF]

Meiners, S.J., Cadotte, M.W., Fridley, J.D., Pickett, S.T.A. and Walker, L.R. 2015. Is successional research nearing its climax? New approaches for understanding dynamic communities. Functional Ecology 29: 154-164. [PDF]

Siefert, A., Lesser, M.R., and Fridley, J.D. 2015. How do climate and dispersal traits limit ranges of tree species along latitudinal and elevational gradients? Global Ecology and Biogeography 24: 581-593. [PDF]


Fridley, J.D. and Sax, D.F. 2014. The imbalance of nature: revisiting a Darwinian framework for invasion biology. Global Ecology and Biogeography 23: 1157-1166. [PDF] [Read coverage of the article in Carl Zimmer's NY Times column.]

Siefert, A., Fridley, J.D., and Ritchie, M.E. 2014. Community functional responses to soil and climate at multiple spatial scales: when does intraspecific variation matter? PLoS ONE 9(10): e111189. [online]

Smith, M.S., Fridley, J.D., Goebel, M., and Bauerle, T.L. 2014. Links between belowground and aboveground resource-related traits reveal species growth strategies that promote invasive advantages. PLoS ONE 9(8): e104189. [online]

Ravenscroft, C.H., Fridley, J.D., and Grime, J.P. 2014. Intraspecific functional differentiation suggests local adaptation to long-term climate change in a calcareous grassland. Journal of Ecology 102: 65-73. [PDF]

Raney, P.A., Fridley, J.D., and Leopold, D.J. 2014. Characterizing microclimate and plant community variation in wetlands. Wetlands 34: 43-53. [PDF]


Heberling, J.M. and Fridley, J.D. 2013. Resource-use strategies of native and invasive plants in Eastern North American forests. New Phytologist 200: 523-533. [PDF]

Fridley, J.D. 2013. Plant invasions across the Northern Hemisphere: a deep-time perspective. Annals of the New York Academy of Sciences 1293: 8-17. [PDF]

Fridley, J.D. 2013. Successional convergence, stochastic assembly, and the future of tropical forests. Journal of Vegetation Science 24: 415-416. [PDF]

Smith, M.S., Fridley, J.D., Yin, J., and Bauerle, T.L. 2013. Contrasting xylem vessel constraints on hydraulic conductivity between native and non-native woody understory species. Frontiers in Plant Science 4: 486. [PDF]

Eallonardo, A.S., Leopold, D.J., Fridley, J.D., and Stella, J.C. 2013. Salinity tolerance and the decoupling of resource axis plant traits. Journal of Vegetation Science 24: 365-274. [PDF]


Fridley, J.D. 2012. Extended leaf phenology and the autumn niche in deciduous forest invasions. Nature 485: 359-362. [PDF]

Fridley, J.D., Grime, J.P., Huston, M.A., Pierce, S., Smart, S.M., Thompson, K., Börger, L., Brooker, R.W., Gross, N., Liancourt, P., Michalet, R. and Le Bagousse-Pinguet, Y. 2012. Comment on "Productivity is a poor predictor of plant species richness". Science 335: 1441b. [online]

Heberling, J.M. and Fridley, J.D. 2012. Biogeographic constraints on the worldwide leaf economics spectrum. Global Ecology and Biogeography 21:1137-1146. [PDF]

Siefert, A., Ravenscoft, C., Althoff, D., Alvarez-Yapiz, J.C., Carter, E., Glennon, K., Heberling, M., Jo, I., Pontes, A., Sauer, A., Willis, A., and Fridley, J.D. 2012. Scale dependence of vegetation-environment relationships: a meta-analysis of multivariate data. Journal of Vegetation Science 23: 942-951. [PDF]

Fridley, J.D. and Wright, J.P. 2012. Drivers of secondary succession rates across temperate latitudes of the Eastern U.S.: climate, soils, and species pools. Oecologia 168: 1069-1077. [PDF]

Grime, J.P. and Fridley, J.D. 2012. Species richness in herbaceous vegetation. In: Levin, S.A., Ed. Encyclopedia of Biodiversity, 2nd Edition. Elsevier.


Askew, A.P., Fridley, J.D., and Grime, J.P. 2011. Predicting future impacts of global change on grassland ecosystems: the role of long-term manipulations of climate. In Lemaire, G. (ed), Grassland Productivity and Ecosystem Services. CABI.

Moser, B., Fridley, J.D., Grime, J.P., and Askew, A.P. 2011. Simulated migration in a long-term climate change experiment: invasions impeded by dispersal limitation, not biotic resistance. Journal of Ecology 99: 1229-1236. [PDF]

Fridley, J.D., Grime, J.P., Askew, A.P., Moser, B., and Stevens, C.J. 2011. Soil heterogeneity buffers community response to climate change in species-rich grassland. Global Change Biology 17: 2002-2011. [PDF]

Manthey, M., Fridley, J.D. and Peet, R.K. 2011. Niche expansion after competitor extinction? A comparative assessment of habitat generalists and specialists in the tree floras of southeastern North America and southeastern Europe. Journal of Biogeography 38: 840-853. [PDF]

Fridley, J.D. 2011. Biodiversity as a bulwark against invasion: conceptual threads since Elton. In: Richardson, D.M. ed. Fifty years of invasion ecology: the legacy of Charles Elton. Blackwell, pp. 121-130.

Fridley, J.D. 2011. Invasibility of communities and ecosystems. In: Simberloff, D. and Rejmanek, M., eds. Encyclopedia of biological invasions. University of California Press, pp. 356-360. [PDF]


Frank, D.A., Pontes, A.W., Maine, E.M., Caruana, J., Raina, R., Raina, S., and Fridley, J.D. 2010. Grassland root communities: species distributions and how they are linked to aboveground abundance. Ecology 91: 3201-3209. [PDF]

Fridley, J.D. and Grime, J.P. 2010. Community- and ecosystem-level consequences of intraspecific genetic diversity in grassland microcosms of varying species diversity. Ecology 91: 2272-2283. [PDF]

Wright, J.P. and Fridley, J.D. 2010. Biogeographic synthesis of secondary succession rates in Eastern North America. Journal of Biogeography 37: 1584-1596. [PDF] (Editor's Choice Sept 2010)


Fridley, J.D., Senft, A., and Peet, R.K. 2009. Vegetation structure of field margins and adjacent forests in agricultural landscapes of the North Carolina Piedmont (USA). Castanea 74: 327-339. [PDF]

Fridley, J.D. 2009. Downscaling climate over complex terrain: high fine-scale spatial variation of near-ground temperatures in a montane forested landscape (Great Smoky Mountains, USA). Journal of Applied Meteorology and Climatology 48: 1033-1049. [PDF] [Supplemental Information]

Manthey, M. and Fridley, J.D. 2009. Beta diversity metrics and the estimation of niche width via species co-occurrence data: reply to Zeleny. Journal of Ecology 97: 18-22. [PDF]


Fridley, J.D. 2008. Of Asian forests and European fields: Eastern U.S. plant invasions in a global floristic context. PLoS ONE 3: e3630. [online] [PDF]

Grime, J.P., Fridley, J.D., Askew, A.P., Thompson, K., Hodgson, J.G., and Bennett, C.R. 2008. Long-term resistance to simulated climate change in an infertile grassland. Proceedings of the National Academy of Sciences USA 105: 10028-10032. [PDF]

Palmer, M., McGlinn, D., and Fridley, J.D. 2008. Artifacts and artifictions in biodiversity research. Folia Geobotanica 43: 245-257. [PDF]


Qian, H., Fridley, J.D., and Palmer, M. W. 2007. The latitudinal gradient of species-area relationships of vascular plants of North America. The American Naturalist 170: 690-701. [PDF]

Fridley, J.D., Grime, J.P., and Bilton, M. 2007. Genetic identity of interspecific neighbours mediates plant responses to competition and environmental variation in a species-rich grassland. Journal of Ecology 95: 908-915. [PDF] (Editor's Choice of the Sept issue)

Fridley, J.D., Vandermast, D.B., Kuppinger, D.M., Manthey, M., and Peet, R.K. 2007. Co-occurrence-based assessment of habitat generalists and specialists: a new approach for the measurement of niche width. Journal of Ecology 95: 707-722. [PDF]

Fridley, J.D., Stachowicz, J.J., Naeem, S., Sax, D.F., Seabloom, E.W., Smith, M.D., Stohlgren, T.J., Tilman, D., and Von Holle, B. 2007. The invasion paradox: reconciling pattern and process in species invasions. Ecology 88: 3-17. [PDF]

to 2006

Fridley, J.D., Peet, R.K., van der Maarel, E., and Willems, J.H. 2006. Integration of local and regional species-area relationships from space-time species accumulation. The American Naturalist 168:133-143. [PDF]

Fridley, J.D., Qian, H., White, P.S., and Palmer, M.W. 2006. Plant species invasions along the latitudinal gradient in the United States: comment. Ecology 87:3209-3213. [PDF]

Bruno, J.F., Fridley, J.D., Bromberg, K.D., and Bertness, M.D. 2005. Insights into biotic interactions from studies of species invasions. In: Sax, D.F., Gaines, S.D., and Stachowicz, J.J., eds. Species invasions: insights into ecology, evolution, and biogeography. Sinauer, Sunderland, MA.

Fridley, J.D., Peet, R.K., White, PS, and Wentworth, T.R. 2005. Connecting fine- and broad-scale species-area relationships of Southeastern U.S. flora. Ecology 86: 1172-1177. [PDF]

Fridley, J.D., Brown, R.L., and Bruno, J.F. 2004. Null models of exotic invasion and scale-dependent patterns of native and exotic species richness. Ecology 85: 3215-3222. [PDF]

Peet, R.K., Fridley, J.D., and Gramling, J.M. 2003. Variation in species richness and species pool size across a pH gradient in forests of the Southern Blue Ridge Mountains. Folia Geobotanica 38: 391-401. [PDF]

Fridley, J.D. 2003. Diversity effects on productivity in different light and nutrient environments: an experiment with communities of annual plants. Journal of Ecology 91: 396-406. [PDF]

Brown, R.L. and Fridley, J.D. 2003. Control of plant species diversity and community invasibility by species immigration: seed richness versus seed density. Oikos 102: 15-24. [PDF]

Fridley, J.D. 2002. Resource availability dominates and alters the relationship between species diversity and ecosystem productivity in experimental plant communities. Oecologia 132: 271-277. [PDF]

Fridley, J.D. 2001. The influence of species diversity on ecosystem productivity: how, where, and why? Oikos 93: 514-526. [PDF]

Huston, M.A., Aarssen, L.W., Austin, M.P., Cade, B.S., Fridley, J.D., Garnier, E., Grime, J.P., Hodgson, J., Lauenroth, W.K., Thompson, K., Vandermeer, J.H., and Wardle, D.A. 2000. No consistent effect of plant diversity on productivity (Technical Comment). Science 289: 1255a. [PDF]

Fridley, J.D. 1998. The flora and fauna of Duke Paul's Missouri. In: Hans von Sachsen-Altenburg and Robert L. Dyer, Duke Paul of Wuerttemberg on the Missouri Frontier: 1823, 1830 and 1851. Pekitanoui, Boonville, MO.